Defensive lineman John Franklin-Myers (knee) and tight end Evan Engram (calf) improved to full participants on Thursday.

Inside linebacker Alex Singleton (thumb) was listed as a full participant.

Linebacker Dre Greenlaw (quad) and defensive tackle Malcolm Roach (calf) were among the players who were ruled out for Sunday's season opener against the Titans.

Roach (calf) and Tillman (hamstring) were both listed as limited participants.

Tight end Nate Adkins (ankle) and linebacker Dre Greenlaw (quad) were the lone Broncos to not participate in practice on Wednesday.

"I can't say he's going to be medically ready to play Week 1, but close enough to where we're probably going to hold him [on the active roster]," Head Coach Sean Payton said Thursday.

Linebacker Dre Greenlaw also saw increased action on Tuesday.

Head Coach Sean Payton also said tackle Garett Bolles, who exited practice early on Thursday, is "going to be fine." Bolles had an asthma-related issue, according to Payton.

The Broncos expect to regain a pair of defenders at practice next week.

Head Coach Sean Payton said Singleton was expected to undergo surgery on Tuesday before returning to Denver on Wednesday.

"The news for us was good news, in that it's a ligament in his foot and it's not a tendon," Head Coach Sean Payton said.