With injuries striking players such as Jake Butt, Drew Lock and Theo Riddick in recent weeks, the Broncos had some tricky waters to navigate and will likely be busy in the coming days.

Malik Reed was a force, Khalfani Muhammad and the running game got going in the third quarter, and punter Colby Wadman had an impressive night as the Broncos concluded the preseason.

Keishawn Bierria's fourth-and-goal tackle was one of two fourth-down stops in the red zone that defined an outstanding night for the Broncos' front-seven depth.

QBs Kevin Hogan and Brett Rypien are expected to get the snaps in a game that will likely showcase players competing to provide roster depth.

Emmanuel Sanders looked like his usual, explosive self. Given how the Broncos need his vertical speed, that was the best news of the night.

There are not many first-team positions up for grabs, but the returner spots are. Kelvin McKnight, who had an 18-yard punt return in the preseason opener, is in the mix.

Drew Lock noticed a difference in the energy of Denver's offense, which led to a productive practice. "We got the juices flowing early, so whenever that happens, you know it's going to be a good day," he said.

The 49ers ratcheted up the intensity, which players like Chris Harris Jr. embraced so they could learn more about their team. "I want to see who wants to compete, who shies away, who doesn't like the competition," Harris said.

But with San Francisco's George Kittle dealing with a calf injury, Chris Harris Jr. and the Broncos may not get the challenge they hoped to receive in the upcoming joint practices.

Now "it's time to get serious," as wide receiver Tim Patrick said. But on Tuesday, they treated the fans to an aerial circus.

Previously a safety, Carter now finds himself learning how to be an inside linebacker.

As the only healthy fullback on the roster, he will be busy with the offense during its team periods. But his work on special teams could determine whether he makes the 53-man roster.